Can only humans have knowledge?

Is it only humans that can possess knowledge? There's a widely held view that computers can't have knowledge: that a computer can only hold information, while it is the humans who programmed the computer who have the knowledge. The AKI model of David Williams correctly, in my view, explicitly ties knowledge to action. Any system that can perform autonomous actions in response to environmental cues is knowledgeable. The structures (biological, mechanical, and/or electronic) that determine responses are its knowledge.